reference: Csuzdi, Cs. 1995: A catalogue of Benhamiinae species (Annelida: Oligochaeta: Acanthodrilidae).- Ann Naturhist. Mus. Wien, 97B:99-123.

 

original name

valid name
adjelana Michaelsen, 1915; Dichogaster, Dichogaster (Diplothecodrilus) adjelana Michaelsen, 1915
aequalis Michaelsen, 1914; Dichogaster spec. inc. sed.
affinis Michaelsen, l890; Benhamia Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
africana Omodeo, 1955; Neogaster

Pickfordia (Omodeoscolex) africana (Omodeo, 1955)

agilis Omodeo & Vaillaud, 1967; Dichogaster Dichogaster (Dichogaster) wenkei (Michaelsen, 1931)
aidae Righi, 1975; Neogaster Neogaster aidae Righi, 1975
aloysii-sabaudiae Cognetti, 1906; Dichogaster Dichogaster (Diplothecodrilus) aloysiisabaudiae Cognetti, 1906
americanus Cernosvitov, 1934; Neogaster Neogaster americana Cernosvitov, 1934
andina Cognetti, 1904; Dichogaster Dichogaster (Diplothecodrilus) andina Cognetti, 1904
andina evae Righi, Ayres & Bittencourt, 1978; Dichogaster Dichogaster (Diplothecodrilus) andina Cognetti, 1904
angeloi Righi, 1988; Neogaster Neogaster angeloi Righi, 1988
anguana Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) anguana Michaelsen, 1915
annae Horst, 1893; Benhamia Dichogaster (Diplothecodrilus) annae (Horst, 1893)
anomala Omodeo, 1955; Millsonia Millsonia anomala Omodeo, 1955
anomala f. leptocystis Omodeo & Vaillaud, 1967; Millsonia Millsonia omodeoi Sims, 1986
arborea James, 1996; Dichogaster Eutrigaster (Graffia) arborea (James, 1996)
arboricola Wasawo & Omodeo, 1963; Dichogaster Dichogaster (Dichogaster) arboricola Wasawo & Omodeo, 1963
arcifera Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) arcifera Omodeo, 1958
artesetosa Sims, 1986; Millsonia Millsonia artesetosa Sims, 1986
ashantiensis Sims, 1986; Millsonia Millsonia ashantiensis Sims, 1986
asymmetricus Csuzdi, 1995; Monothecodrilus Monothecodrilus asymmetricus Csuzdi, 1995
athena James, 1996; Dichogaster Eutrigaster (Eutrigaster) athena (James, 1996)
austeni Beddard, 1901; Benhamia Dichogaster (Diplothecodrilus) austeni (Beddard, 1901)
baeri Sciacchitano, 1952; Dichogaster Dichogaster (Dichogaster) baeri Sciacchitano, 1952
badajos Righi, Ayres & Bittencourt, 1978; Dichogaster Dichogaster (Diplothecodrilus) badajos Righi & Al., 1978
bagiloana Michaelsen, 1937; Dichogaster Dichogaster (Diplothecodrilus) bagiloana Michaelsen, 1937
balantina Omodeo, 1958; Benhamia Benhamiona balantina (Omodeo, 1958)
balantina Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) balantina Michaelsen, 1915
basseterrensis James, 1996; Dichogaster Eutrigaster (Graffia) basseterrensis James, 1996
bataguensis Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) aloysiisabaudiae Cognetti, 1906
baumanni Michaelsen, 1897; Benhamia Benhamiona baumanni (Michaelsen, 1897)
beauforti Michaelsen, 1933; Wegeneriella Wegeneriona beauforti Michaelsen, 1933
beddardi Horst, 1888; Acanthodrilus Benhamiona beddardi (Horst, 1888)
belenensis Righi, 1988; Wegeneriona Wegeneriona belenensis Righi, 1988
bidjumensis Michaelsen, 1915; Monogaster Monogaster bidjumensis Michaelsen, 1915
biggei Michaelsen, 1915; Dichogaster Benhamiona biggei (Michaelsen, 1915)
bolamensis Cognetti, 1910; Dichogaster Guineoscolex bolamensis (Cognetti, 1910)
bolaui Michaelsen, 1891; Benhamia Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
bolaui amphibiotica Dahl, 1957; Dichogaster Dichogaster (Diplothecodrilus) amphibiotica Dahl, 1957
bolaui v. decanephra Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
bolaui v. malabaricus Stephenson, 1920; Dichogaster Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
bolaui pacifica Eisen, 1900; Benhamia Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
brasiliana Cernosvitov, 1939; Wegeneriona Wegeneriona brasiliana Cernosvitov, 1939
braunsi Michaelsen, 1895; ? Microdrilus Omodeona braunsi (Michaelsen, 1895)
brevicingulata Sims, 1986; Millsonia Millsonia brevicingulata Sims, 1986
budgetti Beddard, 1900; Benhamia Benhamiona budgetti (Beddard, 1900)
büttikoferii Horst, 1884; Acanthodrilus Benhamiona buettikoferii (Horst, 1884)
büttneri Michaelsen, 1892; Benhamia Dichogaster (Dichogaster) buettneri (Michaelsen, 1892)
caecifera Benham, 1894; Benhamia Millsonia caecifera (Benham, 1894)
caesiticifusca James, 1996 Dichogaster Eutrigaster (Graffia) caesiticifusca (James, 1996)
cagnii Cognetti, 1906; Dichogaster Dichogaster (Diplothecodrilus) cagnii Cognetti, 1906
callaina James, 1996; Dichogaster Eutrigaster (Graffia) callaina ( James, 1996)
candida Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) candida Omodeo, 1958
capilliseta Omodeo, 1958; Benhamia Benhamiona capilliseta (Omodeo, 1958)
castanea Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) castanea (Michaelsen, 1896)
chapini Michaelsen, l936; Dichogaster Dichogaster (Diplothecodrilus) chapini Michaelsen, 1936
chappuisi Cernosvitov, 1938; Dichogaster Dichogaster (Diplothecodrilus) heterochaeta Michaelsen, 1922
centralis Sims, 1986; Millsonia Millsonia centralis Sims, 1986
cernosvitovi Righi & Caballero, 1970; Wegeneriona Wegeneriona cernosvitovi Righi & Caballero, 1970
cheranganiensis Cernosvitov, 1938; Dichogaster Dichogaster (Diplothecodrilus) annae (Horst, 1893)
christyana Michaelsen, 1937; Dichogaster Dichogaster (Diplothecodrilus) christyana Michaelsen, 1937
cisatlantica Omodeo, 1955; Neogaster Pickfordia (Omodeoscolex) cisatlantica (Omodeo, 1955)
complanata Michaelsen, 1897; Benhamia Dichogaster (Diplothecodrilus) complanata (Michaelsen, 1897)
congica Horst, 1894; Benhamia Dichogaster (Diplothecodrilus) congica (Horst, 1894)
congica Zicsi & Csuzdi 1989; Wegeneriella Wegeneriella congica Zicsi & Csuzdi, 1989
coronata Omodeo, 1958; Benhamia Guineoscolex coronatus (Omodeo, 1958)
corticis Michaelsen, 1899; Benhamia Dichogaster (Dichogaster) corticis (Michaelsen, 1899)
crassa Beddard, 1893; Benhamia Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
crateris Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) crateris Michaelsen, 1912
crawi Eisen, 1900; Dichogaster Dichogaster (Diplothecodrilus) saliens (Beddard, 1892)
cruciventris Sims, 1986; Millsonia Millsonia cruciventris Sims, 1986
culminis Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) culminis (Michaelsen, 1896)
cultrifera Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) cultrifera Michaelsen, 1903
curgensis Michaelsen, 1921; Dichogaster Dichogaster (Diplothecodrilus) annae (Horst, 1893)
curgensis v. unilocularis Stephenson, 1931; Dichogaster Dichogaster (Diplothecodrilus) annae (Horst, 1893)
curta Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) curta (Michaelsen, 1896)
curvisetifera Csuzdi & Zicsi, 1994; Dichogaster Dichogaster (Diplothecodrilus) curvisetifera Csuzdi & Zicsi, 1994
cyrtochaeta Cognetti, 1910; Dichogaster Dichogaster (Dichogaster) cyrtochaeta Cognetti, 1910
daemoniaca Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) daemoniaca Cognetti, 1907
damonis Beddard, 1888; Dichogaster Dichogaster (Dichogaster) damonis
decorata Graff, 1957; Dichogaster Eutrigaster (Eutrigaster) vialis (Michaelsen, 1912)
deflecticista Michaelsen, 1935; Dichogaster Dichogaster (Diplothecodrilus) deflecticista Michaelsen, 1935
dewittei Csuzdi & Zicsi , 1994; Dichogaster Dichogaster (Diplothecodrilus) dewittei Csuzdi & Zicsi, 1994
ditheca Sims, 1965; Millsonia Beddard, 1894 Millsonia ditheca Sims, 1965
ditheca Omodeo, 1958; Pickfordia Pickfordia (Pickfordia) ditheca Omodeo, 1958
divergens Cognetti, 1905; Notiodrilus Pickfordia (Omodeoscolex) d. divergens (Cognetti, 1905)
divergens itapecu Righi, et Al.; Wegeneriella Pickfordia (Omodeoscolex) d. itapecu (Righi & Al., 1978)
djemdjemensis Stephenson, 1932; Dichogaster Dichogaster (Diplothecodrilus) djemjemensis Stephenson, 1932
dokoensis Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) dokoensis Michaelsen, 1903
dominicae Lavelle, 1981; Agastrodrilus Agastrodrilus dominicae Lavelle, 1981
dorsalis Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) culminis (Michaelsen, 1896)
doveri Stephenson, 1930; Dichogaster Dichogaster (Diplothecodrilus) modiglianii (Rosa, 1896)
dudichi Csuzdi, 1995; Monothecodrilus Monothecodrilus dudichi Csuzdi, 1995
duwonica Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) culminis (Michaelsen, 1896)
dzwilloi Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) dzwilloi Csuzdi & Zicsi, 1989
ehrhardti Michaelsen, 1898; Balanta Dichogaster (Dichogaster) ehrhardti (Michaelsen, 1898)
ehrhardti v. linnelli Michaelsen, 1910; Dichogaster Dichogaster (Dichogaster) ehrhardti (Michaelsen, 1897)
eiseni Csuzdi & Zicsi, 1991; Eutrigaster (Graffia) Eutrigaster (Graffia) eiseni Csuzdi & Zicsi, 1991
elgonensis Michaelsen, 1937; Dichogaster Dichogaster (Diplothecodrilus) elgonensis Michaelsen, 1937
equatorialis Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) aequatorialis (Michaelsen, 1896)
enchytraeus Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) enchytraeus Omodeo, 1958
erlangeri Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) erlangeri Michaelsen, 1903
ernesti Michaelsen, 1897; Benhamia Dichogaster (Diplothecodrilus) ernesti (Michaelsen, 1897)
esca Stephenson, 1931; Dichogaster Benhamiona esca (Stephenson, 1931)
eudrilina Cognetti, 1909; Dichogaster Millsonia nigra Beddard, 1894
excelsa Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) excelsa Cognetti, 1907
fallax Cognetti, 1910; Dichogaster Omodeona fallax (Cognetti, 1910)
feae Cognetti, 1910; Dichogaster Dichogaster (Dichogaster) feai Cognetti, 1910
ficta Michaelsen, 1934; Dichogaster Dichogaster (Diplothecodrilus) congica (Horst,1894)
flagellifera Stephenson, 1933; Dichogaster Dichogaster (Diplothecodrilus) flagellifera Stephenson, 1933
flandria Michaelsen, 1936; Dichogaster Dichogaster (Diplothecodrilus) flandria Michaelsen, 1936
floresiana Horst, 1893; Benhamia Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
foetida Michaelsen, 1932; Dichogaster Dichogaster (Diplothecodrilus) foetida Michaelsen, 1932
fouoriensis Csuzdi, 1996; Dichogaster (Dichogaster) Dichogaster (Dichogaster) fouoriensis Csuzdi, 1996
franzi Csuzdi & Zicsi, 1991; Eutrigaster (Eutrigaster) Eutrigaster (Eutrigaster) montecyanensis Sims, 1987,
frickei Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) frickei Michaelsen, 1912
fula Sims, 1967; Benhamia Guineoscolex bolamensis (Cognetti, 1910)
gabunensis Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) gabunensis Michaelsen, 1915
gagzoi Michaelsen, 1908; Dichogaster Eutrigaster (Graffia) gagzoi (Michaelsen, 1908)
gambiana Beddard, 1902; Benhamia Benhamiona gambiana (Beddard, 1902)
gardullaensis Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) gardullaensis Michaelsen, 1903
gatesi Csuzdi, 1996; Dichogaster (Diplothecodrilus) Dichogaster (Diplothecodrilus) gatesi Csuzdi, 1996
gavrilovi Righi & Caballero, 1970; Neogaster Neogaster gavrilovi Righi & Caballero, 1970
gestri Cognetti, 1910; Dichogaster Dichogaster (Dichogaster) gestroi Cognetti, 1910
ghanensis Sims, 1965; Millsonia Millsonia ghanensis Sims, 1965
girija James, 1996; Dichogaster Eutrigaster (Graffia) girija (James, 1996)
godeffroyi Michaelsen, 1890; Benhamia Eutrigaster (Graffia) godeffroyi (Michaelsen, 1890)
gofaensis Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) gofaensis Michaelsen, 1903
golaensis Michaelsen, 1913; Dichogaster Dichogaster (Dichogaster) golaensis Michaelsen, 1913
gracilis Michaelsen, 1892; Benhamia Dichogaster (Diplothecodrilus) gracilis (Michaelsen, 1892)
gracilis v. metandra Omodeo, 1973; Dichogaster Dichogaster (Diplothecodrilus) gracilis (Michaelsen, 1892)
graffi Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Dichogaster) graffi Csuzdi & Zicsi, 1989
grandis Sims, 1987; Eutrigaster Eutrigaster (Eutrigaster) grandis Sims, 1987
greeffi Michaelsen, 1902; Dichogaster Dichogaster (Dichogaster) greeffi, Michaelsen, 1902
griffini Csuzdi, 1995; Guineoscolex Guineoscolex griffini Csuzdi, 1995
guadeloupensis James, 1996; Dichogaster Eutrigaster (Graffia) guadeloupensis ( James, 1996)
guatemalae Eisen, 1900; Benhamia Eutrigaster (Graffia) guatemalae (Eisen, 1900)
guetare Righi & Merino, 1987; Dichogaster Eutrigaster (Eutrigaster) guetare (Righi & Merino, 1987)
guildingi Baird, 1871; Lumbricus Dichogaster (Diplothecodrilus) guildingi (Baird, 1871)
hamatus Stephenson, 1932; Dichogaster Dichogaster (Diplothecodrilus) scotti Stephenson, 1932
hamburgensis Omodeo, 1958; Dichogaster Dichogaster (Diplothecodrilus) kigogoana Stephenson, 1933
hammon Somon, 1995; Dichogaster Dichogaster (Dichogaster) hammon Somon, 1995
hansi Michaelsen, 1914; Dichogaster Benhamiona hansi (Michaelsen, 1914)
hartmeyeri Michaelsen, 1908; Dichogaster Eutrigaster (Graffia) hartmeyeri (Michaelsen, 1908)
hatomaana Ohfuchi, 1957; Dichogaster Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
hemiandra Omodeo, 1973; Dichogaster Dichogaster (Diplothecodrilus) hemiandra Omodeo, 1973
hemibalantina Omodeo, 1958; Pickfordia Pickfordia (Pickfordia) hemibalantina Omodeo, 1958
hemina Sims, 1965; Millsonia Millsonia hemina Sims, 1965
heterochaeta Michaelsen, 1922; Dichogaster Dichogaster (Diplothecodrilus) heterochaeta Michaelsen, 1922
heteronephra Michaelsen, 1897; Benhamia Millsonia heteronephra (Michaelsen, 1897)
hilaris Cognetti, 1904; Dichogaster Eutrigaster (Eutrigaster) hilaris (Cognetti, 1904)
horsti Michaelsen, 1898; Benhamia Benhamiona horsti (Michaelsen, 1898)
hortensis Sims, 1986; Millsonia Beddard, 1894 Millsonia hortensis Sims, 1986
hupferi Michaelsen, 1891; Dichogaster Benhamiona hupferi (Michaelsen, 1891)
ibaia Righi, Ayres & Bittencourt, 1978; Dichogaster Dichogaster (Diplothecodrilus) ibaia Righi & Al., 1978
inaequalis Michaelsen, 1914; Dichogaster Guineoscolex inaequalis (Michaelsen, 1914)
inaequalis f. nimbai Omodeo, 1958; Benhamia Guineoscolex inaequalis (Michaelsen, 1914)
inermis Michaelsen, 1892; Benhamia Millsonia inermis (Michaelsen, 1892)
inermis v. guttata Michaelsen, 1913; Dichogaster Millsonia guttata (Michaelsen, 1913)
insignis Michaelsen, 1922; Dichogaster spec. inc. sed.
insolitus Sims, 1986; Agastrodrilus Agastrodrilus insolitus Sims, 1986
insularis Michaelsen, 1895; Benhamia Dichogaster (Dichogaster) insularis (Michaelsen, 1895)
insulsa Cognetti, 1910; Dichogaster Dichogaster (Diplothecodrilus) insulsa Cognetti, 1910
intermedia Michaelsen, 1891; Benhamia Dichogaster (Diplothecodrilus) intermedia (Michaelsen, 1891)
ireneae Csuzdi, 1996; Dichogaster (Diplothecodrilus) Dichogaster (Diplothecodrilus) ireneae Csuzdi, 1996
itoliensis Michaelsen, 1892; Benhamia Benhamia itoliensis Michaelsen, 1892
itoliensis v. coerulea Michaelsen, 1899; Benhamia Benhamia itoliensis Michaelsen, 1892
itoliensis v. minor Stephenson, 1933; Dichogaster Benhamia itoliensis Michaelsen, 1892
jaculatrix Baylis, 1915; Dichogaster Benhamia itoliensis Michaelsen, 1892
jadwigae Sims, 1986; Millsonia Millsonia jadwigae Sims, 1986
jamaicae Eisen, 1900; Benhamia Dichogaster (Diplothecodrilus) jamaicae (Eisen, 1900)
jamiesoni Csuzdi, 1996; Dichogaster (Diplothecodrilus) Dichogaster (Diplothecodrilus) jamiesoni Csuzdi, 1996
johnstoni Beddard, 1901; Benhamia 1889 Benhamia itoliensis Michaelsen, 1892
jordani Michaelsen, 1937; Dichogaster Dichogaster (Diplothecodrilus) jordani Michaelsen, 1937
kaburomina Michaelsen, 1937; Dichogaster Dichogaster (Diplothecodrilus) kaburomina Michaelsen, 1937
kaffaensis Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) kaffaensis Michaelsen, 1903
kafuruënsis Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) modiglianii (Rosa, 1896)
kakulimana Michaelsen, 1914; Dichogaster spec. inc. sed.
kamerunensis Michaelsen, 1903; Dichogaster spec. inc. sed.
karagoensis Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) karagoensis Michaelsen, 1912
karissimbii Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) karissimbii Michaelsen, 1912
katangae Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) katangae Michaelsen, 1915
keiteli Michaelsen, 1898; Benhamia Eutrigaster (Graffia) keiteli (Michaelsen, 1898)
kepo Righi & Merino, 1987; Dichogaster Eutrigaster (Eutrigaster) kepo (Righi & Merino, 1987)
kenyae Michaelsen, 1914; Dichogaster Dichogaster (Diplothecodrilus) erlangeri Michaelsen, 1903
kigogoana Stephenson, 1933; Dichogaster Dichogaster (Diplothecodrilus) kigogoana Stephenson, 1933
kindiana Michaelsen, 1914; Dichogaster Benhamiona kindiana (Michaelsen, 1914)
kiwuensis Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) kiwuensis Michaelsen, 1912
krugeri Reinecke & Ackermann, 1977; Dichogaster Dichogaster (Diplothecodrilus) krugeri Reinecke & Ackermann, 1977
kunguluensis Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) kungulunensis Michaelsen, 1915
kwidschwiensis Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) kwidschwiensis Michaelsen, 1912
lageniformis Friend, 1916; Dichogaster Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
lamottei Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) lamottei Omodeo, 1958
lamtoiana Omodeo & Vaillaud, 1967; Millsonia Millsonia lamtoiana Omodeo & Vaillaud, 1967
lavellei Sims, 1986; Agastrodrilus Agastrodrilus lavellei Sims, 1986
lavellei Csuzdi, 1992; Dichogaster Dichogaster (Diplothecodrilus) lavellei Csuzdi, 1992
leroyi Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) leroyi Omodeo, 1958
liberiensis Horst, 1895; Benhamia Benhamiona liberiensis (Horst, 1895)
ligula Sims, 1964; Dichogaster Dichogaster (Dichogaster) ligula Sims, 1964
lineri Righi, 1972; Dichogaster Eutrigaster (Eutrigaster) lineri (Righi, 1972)
lituifera Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) lituifera Michaelsen, 1903
loboziana Michaelsen, l915; Dichogaster Dichogaster (Diplothecodrilus) loboziana Michaelsen, 1915
lofaensis Michaelsen, 1914; Dichogaster Dichogaster (Dichogaster) lofaensis Michaelsen, 1914
loveridgei Stephenson, 1933; Dichogaster Dichogaster (Diplothecodrilus) austeni (Beddard, 1901)
lönnbergi Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) loennbergi Michaelsen, 1912
lufirae Michaelsen, 1935; Dichogaster Dichogaster (Diplothecodrilus) lukafuensis Michaelsen, 1935
lukafuensis Michaelsen, 1935; Dichogaster Dichogaster (Diplothecodrilus) lukafuensis Michaelsen, 1935
macfadyeni Csuzdi, 1991; Dichogaster Dichogaster (Diplothecodrilus) macfadyeni Csuzdi, 1991
machadoi Omodeo, 1973; Dichogaster Dichogaster (Diplothecodrilus) machadoi Omodeo, 1973
magnisetosa Omodeo, 1958; Pickfordia (Pickfordia) magnisetosa Omodeo, 1958
majoris Michaelsen, 1914; Dichogaster Dichogaster (Dichogaster) majoris Michaelsen, 1914
malarmata Rosa, 1898; Benhamia Dichogaster (Dichogaster) malarmata (Rosa, 1898)
malayana Horst, 1893; Benhamia Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
mamillata Omodeo, 1958; Benhamia Guineoscolex mamillatus (Omodeo, 1958)
mandinka Sims, 1967; Benhamia Guineoscolex bolamensis (Cognetti, 1910)
mangeri Michaelsen, 1913; Dichogaster Dichogaster (Dichogaster) mangeri Michaelsen, 1913
manni Michaelsen, 1935; Dichogaster Eutrigaster (Graffia) manni (Michaelsen, 1935)
mansfeldi Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) mundamensis (Michaelsen, 1897)
matoubensis James, 1996; Dichogaster Eutrigaster (Graffia) matoubensis (James, 1996)
medellina Michaelsen, 1914; Dichogaster Dichogaster (Diplothecodrilus) medellina Michaelsen, 1914
meridionalis Omodeo, 1973; Millsonia Millsonia meridionalis Omodeo, 1973
mexicana Rosa, 1891; Benhamia 1889 Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
meyaensis Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) meyaensis Csuzdi & Zicsi, 1989
michaelseni Cernosvitov, 1934; Wegeneriella Wegeneriona michaelseni (Cernosvitov, 1934)
michaelseniana Csuzdi & Zicsi,1991; Eutrigaster (Graffia) Eutrigaster (Graffia) michaelseniana Csuzdi & Zicsi., 1991
michaelseni Beddard, 1902; Benhamia Benhamiona michaelseni (Beddard, 1902)
microscolecina Omodeo, 1958; Benhamia Guineoscolex microscolecinus (Omodeo, 1958)
mimus Michaelsen, 1891; Dichogaster Millsonia mima (Michaelsen, 1891)
minuscula Omodeo, 1958; Benhamia Guineoscolex minusculus (Omodeo, 1958)
minutissima Csuzdi, 1996; Dichogaster (Diplothecodrilus) Dichogaster (Diplothecodrilus) minutissima Csuzdi, 1996
misaënsis Michaelsen, 1897; Dichogaster Benhamiona missaensis (Michaelsen, 1897)
moderata Sims, 1986; Millsonia Millsonia moderata Sims, 1986
modesta Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) modesta Michaelsen, 1903
modesta papillifera Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) papillifera Csuzdi & Zicsi, 1989
modiglianii Rosa, 1896; Benhamia Dichogaster (Diplothecodrilus) modiglianii (Rosa, 1896)
mollis Beddard, 1901; Benhamia Benhamia itoliensis Michaelsen, 1892
monotheca Omodeo, 1955; Wegeneriella (Wegeneriella) Wegeneriella monotheca Omodeo, 1955
montana Csuzdi & Zicsi, 1991; Eutrigaster (Graffia) Eutrigaster (Graffia) montana Csuzdi & Zicsi, 1991
montecyanensis Sims, 1987; Eutrigaster Eutrigaster (Eutrigaster) montecyanensis Sims, 1987
monotheca Omodeo, 1958; Benhamia Monothecodrilus monothecus (Omodeo, 1958)
monticellii Cognetti, 1915; Dichogaster Dichogaster (Diplothecodrilus) austeni (Beddard, 1901)
monticola Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) monticola (Michaelsen, 1896)
montis-toi Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) insularis (Michaelsen, 1895)
moorii Beddard, 1901; Benhamia Benhamia itoliensis Michaelsen, 1892
mulataensis Michaelsen, 1903; Dichogaster Dichogaster (Diplothecodrilus) mulataensis Michaelsen, 1903
multivesiculatus Omodeo & Vaillaud, 1967; Agastrodrilus Agastrodrilus multivesiculatus Omodeo & Vaillaud, 1967
mundamensis Michaelsen, 1897; Benhamia Dichogaster (Diplothecodrilus) mundamensis (Michaelsen, 1897)
murithae Csuzdi & Zicsi, 1994; Benhamiona Benhamiona murithae Csuzdi & Zicsi, 1994
musciphila James, 1996; Dichogaster Eutrigaster (Graffia) musciphila (James, 1996)
nana Eisen, 1896; Benhamia Dichogaster (Diplothecodrilus) modiglianii (Rosa,1896)
navana Michaelsen, 1936; Dichogaster Dichogaster (Diplothecodrilus) navana Michaelsen, 1936
nematochaeta Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) nematochaeta Omodeo, 1958
neumanni Michaelsen, 1897; Benhamia Dichogaster (Diplothecodrilus) neumanni (Michaelsen, 1897)
nigra Beddard, 1894; Millsonia Millsonia nigra Beddard, 1894
nilesi Sims, 1986; Millsonia Millsonia nilesi Sims, 1986
nota Sims, 1986; Millsonia Millsonia nota Sims, 1986
notabilis Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) notabilis Omodeo, 1958
nyassana Michaelsen, 1913; Dichogaster Dichogaster (Diplothecodrilus) frickei Michaelsen, 1912
octonephra Rosa, 1895; Benhamia Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
omodeoi Sims, 1986; Millsonia Millsonia omodeoi Sims, 1986
opisthogynus Omodeo & Vaillaud, 1967; Agastrodrilus Agastrodrilus opisthogynus Omodeo & Vaillaud, 1967
oracapensis Sims, 1986; Millsonia Millsonia oracapensis Sims, 1986
oraedivitis Cognetti, 1904; Eutrigaster Eutrigaster (Eutrigaster) oraedivitis Cognetti, 1904
ornata Sciacchitano, 1952; Dichogaster Benhamiona ornata (Sciacchitano, 1952)
orobia Graff, 1957; Dichogaster Eutrigaster (Eutrigaster) orobia (Graff, 1957)
paessleri Michaelsen, 1910; Dichogaster Eutrigaster (Graffia) viridis (Eisen, 1900)
pafuriensis Reinecke & Ackermann, 1977; Dichogaster Dichogaster (Diplothecodrilus) austeni (Beddard, 1901)
pallida Michaelsen, 1892; Benhamia Dichogaster (Diplothecodrilus) gracilis (Michaelsen, 1892)
papillata Eisen, 1896; Benhamia 1889 Dichogaster (Diplothecodrilus) modiglianii (Rosa, 1896)
papillata v. hawaiiensis Eisen, 1900; Benhamia Dichogaster (Diplothecodrilus) modiglianii (Rosa, 1896)
papillosa Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) papillosa Omodeo, 1958
parva Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) annae (Horst, 1893)
pascuali Somon, 1995 Dichogaster Dichogaster (Dichogaster) pascuali Somon, 1995
pauliani Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) pauliani Csuzdi & Zicsi, 1989
penigera Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) penigera Omodeo, 1958
peridiverticulatus Csuzdi, 1995; Guineoscolex Guineoscolex peridiverticulatus Csuzdi, 1995
picadoi Michaelsen, 1912; Dichogaster Eutrigaster (Graffia) picadoi (Michaelsen, 1912)
pinguis Cognetti, 1910; Dichogaster Benhamia pinguis (Cognetti, 1910)
pitahayana Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) tristani Cognetti, 1907
pitti Csuzdi & Zicsi, 1994; Benhamiona Benhamiona pitti Csuzdi & Zicsi, 1994
pobozsnyae Csuzdi, 1994; Eutrigaster (Eutrigaster) Eutrigaster (Eutrigaster) pobozsnyae Csuzdi, 1994
pooensis Cognetti, 1910; Dichogaster Dichogaster (Dichogaster) pooensis Cognetti, 1910
princeps Cognetti, 1910; Dichogaster Dichogaster (Diplothecodrilus) princeps Cognetti, 1910
proandra Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) proandra Omodeo, 1958
proboscoides Sims, 1967; omodeona Omodeona fallax (Cognetti, 1910)
proboscideus Michaelsen, 1897; Dichogaster Dichogaster (Diplothecodrilus) proboscideus Michaelsen, 1897
pseudoplanaria Omodeo, 1958; Pickfordia Pickfordia (Pickfordia) pseudoplanaria Omodeo,1958
pulvillaris Sims, 1986; Millsonia Millsonia pulvillaris Sims, 1986
pumila Sims, 1965; Millsonia Millsonia pumila Sims, 1965
puylaerti Csuzdi & Zicsi, 1994; Benhamiona Benhamiona puylaerti Csuzdi & Zicsi, 1994
reducta Sims, 1967; Benhamia Monothecodrilus reductus (Sims, 1967)
reichardti Michaelsen, 1908; Dichogaster Eutrigaster (Graffia) reichardti (Michaelsen, 1908)
reinckei Michaelsen, 1898; Benhamia Dichogaster (Dichogaster) reinckei (Michaelsen, 1898)
reinckei v. endemica Michaelsen, 1914; Dichogaster Dichogaster (Dichogaster) endemica Michaelsen, 1914
ribaucourti Eisen, 1900; Dichogaster Eutrigaster (Eutrigaster) ribacourti (Eisen, 1900)
riparia Sims, 1986; Millsonia Millsonia riparia Sims, 1986
robertsiana Michaelsen, 1922; Dichogaster Benhamiona robertsiana (Michaelsen, 1922)
roccatii Cognetti, 1906; Dichogaster Dichogaster (Diplothecodrilus) roccatii Cognetti, 1906
rodriguezi Csuzdi, 1994; Eutrigaster (Graffia) Eutrigaster (Graffia) rodriguezi Csuzdi, 1994
rosea Michaelsen, 1889; Benhamia Benhamia rosea Michaelsen, 1889
rosea f. guineana Omodeo, 1958; Benhamia Benhamiona guineana (Omodeo, 1958)
rosea Michaelsen, 1935; Dichogaster Dichogaster (Diplothecodrilus) kigogoana Stephenson, 1933
rubella Michaelsen, 1935; Dichogaster Dichogaster (Diplothecodrilus) rubella Michaelsen, 1935
rubens Beddard, 1894; Millsonia Millsonia mima (Michaelsen, 1891)
rugosa Eisen, 1896; Benhamia Dichogaster (Diplothecodrilus) bolaui (Michaelsen, 1891)
rungweensis Stephenson, 1933; Dichogaster Dichogaster (Diplothecodrilus) rungweensis Stephenson, 1933
ruwenzorii Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) ruwenzorii Cognetti, 1907
saliens Beddard, 1893; Microdrilus Dichogaster (Diplothecodrilus) saliens (Beddard, 1893)
sandoa Michaelsen, 1936; Dichogaster Dichogaster (Diplothecodrilus) stuhlmanni (Michaelsen, 1890)
savanicola Michaelsen, 1915; Dichogaster Dichogaster (Diplothecodrilus) savanicola Michaelsen, 1915
scherbroensis Michaelsen, 1910; Dichogaster Monothecodrilus scherbroensis (Michaelsen, 1910)
schlegeli Horst, 1884; Acanthodrilus Millsonia schlegeli (Horst, 1884)
schomburgki Michaelsen, 1913; Dichogaster Benhamiona schomburgki (Michaelsen, 1913)
schubotzi Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) schubotzi Michaelsen, 1912
scioanus Rosa, 1888; Acanthodrilus spec. inc. sed.
scotti Stephenson, 1932; Dichogaster Dichogaster (Diplothecodrilus) scotti Stephenson, 1932
sellae Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) monticola (Michaelsen, 1896)
septemdecim Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) septemdecim Omodeo, 1958
seriatospina Michaelsen, 1912; Dichogaster Dichogaster (Diplothecodrilus) seriatospina Michaelsen, 1912
servi Righi & Ayres, 1975; Dichogaster Dichogaster (Diplothecodrilus) annae (Horst, 1893)
sexdecim Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) sexdecim Omodeo, 1958
sibitiensis Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Dichogaster) sibitiensis Csuzdi & Zicsi, 1989
silvestrii Michaelsen, 1914; Dichogaster Guineoscolex silvestrii (Michaelsen, 1914)
silvestris Michaelsen, 1896; Benhamia Dichogaster (Diplothecodrilus) silvestris (Michaelsen, 1896)
silvestris cacaois Righi, 1968; Dichogaster Dichogaster (Diplothecodrilus) annae (Horst, 1893)
silvicola Csuzdi, 1996; Loksaia Loksaia silvicola Csuzdi, 1996
singula Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) singula Omodeo, 1958
sinicus Chen, 1938; Dichogaster Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
sinuosus Stephenson, 1931; Dichogaster Dichogaster (Diplothecodrilus) affinis (Michaelsen, 1890)
sokodeana Michaelsen, 1913; Dichogaster Millsonia sokodeana (Michaelsen, 1913)
sorosi Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) sorosi Csuzdi & Zicsi, 1989
spatulifer Csuzdi, 1995; Dudichiodrilus Dudichiodrilus spatulifer Csuzdi, 1995
sporadonephra Cognetti, 1905; Dichogaster Eutrigaster (Graffia) s. sporadonephra (Cognetti, 1905)
sporadonephra disa Righi, 1972; Dichogaster Eutrigaster (Graffia) s. disa (Righi, 1972)
stampflii Horst, 1895; Benhamia spec. inc. sed.
stockhauseni Michaelsen, 1913; Dichogaster Benhamiona stockhauseni (Michaelsen, 1913)
stuhlmanni Michaelsen, 1890; Benhamia Dichogaster (Diplothecodrilus) stuhlmanni (Michaelsen, 1890)
taborana Michaelsen, 1913; Dichogaster Dichogaster (Diplothecodrilus) taborana Michaelsen, 1913
tamiana Cognetti, 1911; Dichogaster Dichogaster (Dichogaster) corticis (Michaelsen, 1899)
tanganyikae Beddard, 1902; Benhamia Dichogaster (Diplothecodrilus) tanganyikae (Beddard, 1902)
tenuis Michaelsen, 1891; Benhamia spec. inc. sed.
tenuiseta Michaelsen, 1936; Dichogaster Dichogaster (Diplothecodrilus) tenuiseta Michaelsen, 1936
terrae-nigrae Omodeo & Vaillaud, 1967; Dichogaster Dichogaster (Dichogaster) terraenigrae Omodeo & Vaillaud, 1967
terricola Michaelsen, 1915; Dichogaster Dichogaster (Dichogaster) terricola Michaelsen, 1915
thibaudi Csuzdi & Zicsi, 1989; Dichogaster Dichogaster (Diplothecodrilus) thibaudi Csuzdi & Zicsi, 1989
thomeana Cognetti, 1910; Dichogaster Dichogaster (Dichogaster) greeffi Michaelsen, 1902
titillata Sims, 1967; Dichogaster Dichogaster (Dichogaster) titillata Sims, 1967
tocaya Righi, et Al., 1978; Wegeneriella Pickfordia (Omodeoscolex) tocaya (Righi & Al., 1978)
togoensis Michaelsen, 1892; Benhamia spec. inc. sed.
toroensis Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) toroensis Cognetti, 1907
townsendi Eisen, 1900; Dichogaster Eutrigaster (Eutrigaster) townsendi (Eisen, 1900)
travancorensis Fedarb, 1898; Benhamia Dichogaster (Diplothecodrilus) annae (Horst, 1893)
tristani Cognetti, 1907; Dichogaster Dichogaster (Diplothecodrilus) tristani Cognetti, 1907
tuzeti Omodeo, 1955; Dichogaster Dichogaster (Dichogaster) tuzeti Omodeo, 1955
ufipana Michaelsen, 1913; Dichogaster Dichogaster (Diplothecodrilus) ufipana Michaelsen, 1913
uhleri Michaelsen, 1935; Dichogaster Eutrigaster (Graffia) uhleri (Michaelsen, 1935)
undecim Omodeo, 1958; Dichogaster Dichogaster (Dichogaster) undecim Omodeo, 1958
valdiviae Michaelsen, 1903; ? Notiodrilus Wegeneriella valdiviae (Michaelsen, 1903)
variabilis Cernosvitov, 1938; Dichogaster Dichogaster (Diplothecodrilus) aequatorialis (Michaelsen, 1896)
verens Cognetti, 1905; Dichogaster Eutrigaster (Graffia) verens (Cognetti, 1905)
vialis Michaelsen, 1912; Dichogaster Eutrigaster (Eutrigaster) vialis (Michaelsen, 1912)
viridis Eisen, 1900; Benhamia Eutrigaster (Graffia) viridis (Eisen, 1900)
wangaensis Michaelsen, 1905; Dichogaster Dichogaster (Diplothecodrilus) wangaensis Michaelsen, 1905
wenkei Michaelsen, 1931; Balanta Dichogaster (Dichogaster) wenkei (Michaelsen, 1931)
whytei Beddard, 1893; Benhamia spec. inc. sed.

reference: Csuzdi, Cs. 1995: A catalogue of Benhamiinae species (Annelida: Oligochaeta: Acanthodrilidae).- Ann Naturhist. Mus. Wien, 97B:99-123.

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